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Epistatic interactions between genetic disorders of hemoglobin can explain why the sickle-cell gene is uncommon in the Mediterranean
Penman, Bridget S..  Proceedings of the National Academy of Sciences of the United States of America.  v. 106. no. 50.   2009-12-01. p. 21242 - 21246... issn: 0027-8424 .   

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